Publications

Current preprints

• Pohle, A., Hoffmann, R., Nützel, A., Seuss, B., Aubrechtová, M., Kröger, B., Stevens, K. and Immenhauser, A. (2025). Microstructural and geochemical evidence offers a solution to the cephalopod cameral deposits riddle. bioRxiv. https://doi.org/10.1101/2025.02.25.639475.
• Pohle, A., Stevens, K., Hoffmann, R., and Immenhauser, A. (2024). Phylogeochemistry: exploring evolutionary constraints on belemnite rostrum element composition. EGUsphere. https://doi.org/10.5194/egusphere-2024-3383.

Peer-reviewed articles

2025

• Klug, C., Pohle, A., Bolliger, T., Hagdorn, H., & Lehmann, J. (2025). Taphonomic and reworking processes isolating cephalopod septa and chamber fillings. Lethaia, 58, 1-18. https://doi.org/10.18261/let.58.2.1

2024

• Klug, C., Schweigert, G., Hoffmann, R., Fuchs, D., Pohle, A., Weis, R. & De Baets, K. (2024). Anatomy and size of Megateuthis, the largest belemnite. Swiss Journal of Palaeontology, 143, 23. https://doi.org/10.1186/s13358-024-00320-x
• Pohle, A. & Klug, C. (2024). Orthoceratoid and coleoid cephalopods from the Middle Triassic of Switzerland with an updated taxonomic framework for Triassic Orthoceratoidea. Swiss Journal of Palaeontology, 143, 14. https://doi.org/10.1186/s13358-024-00307-8
• Pohle, A., Jell, P. & Klug, C. (2024). Plectronoceratids (Cephalopoda) from the latest Cambrian at Black Mountain, Queensland, reveal complex three-dimensional siphuncle morphology, with major taxonomic implications. PeerJ, 12, e17003. https://doi.org/10.7717/peerj.17003

2023

• Cichowolski, M., Vaccari, N., Pohle, A., Morón-Alfonso, D.A., Vaucher, R. & Waisfeld, B.G. (2023). Early Tremadocian cephalopods from Santa Rosita Formation in NW Argentina: the oldest record for South America. Acta Palaeontologica Polonica 68(4), 583-601. https://doi.org/10.4202/app.01103.2023
• Barido-Sottani, J., Pohle, A., De Baets, K., Murdock, D. & Warnock, R.C.M. (2023). Putting the F into FBD analysis: tree constraints or morphological data? Palaeontology, 66(6), e12679. https://doi.org/10.1111/pala.12679
• Klug, C., Hoffmann, R., Tischlinger, H., Fuchs, D., Pohle, A., Rowe, A., et al. (2023). ‘Arm brains’ (axial nerves) of Jurassic coleoids and the evolution of coleoid neuroanatomy. Swiss Journal of Palaeontology, 142, 22. https://doi.org/10.1186/s13358-023-00285-3
• Stevens, K.*, Pohle, A.*, Hoffmann, R., & Immenhauser, A. (2023). Bayesian inference reveals a complex evolutionary history of belemnites. Palaeontologia Electronica, 26(1), a13. https://doi.org/10.26879/1239
• Klug, C., Coates, M., Frey, M. Greif, M., Jobbins, M., Pohle, A., Lagnaoui, A., Bel Haouz, W. & Ginter, M. (2023). Broad snouted cladoselachian with sensory specialization at the base of modern chondrichthyans. Swiss Journal of Palaeontology, 142, 2. https://doi.org/10.1186/s13358-023-00266-6

*Contributed equally

2022

• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Early cephalopod evolution clarified through Bayesian phylogenetic inference. BMC Biology, 20, 88. https://doi.org/10.1186/s12915-022-01284-5

2021

• Kröger, B. & Pohle, A. (2021). Early-Middle Ordovician cephalopods from Ny Friesland, Spitsbergen - a pelagic fauna with Laurentian affinities. European Journal of Taxonomy, 783(1), 1–102. https://doi.org/10.5852/ejt.2021.783.1601
• Fang, X., Pohle, A., Kröger, B., Aubrechtová, M., Burrett, C., Zhang, Y. & Zhang, Y. (2021). Phylogeny of Middle-Late Ordovician lituitid cephalopods based on cladistic analysis. Journal of Systematic Palaeontology, 19(9), 633–650. https://doi.org/10.1080/14772019.2021.1944354
• Klug, C., Pohle, A., Roth, R., Hoffmann, R., Wani, R. & Tajika, A. (2021). Preservation of nautilid soft parts inside and outside the conch interpreted as central nervous system, eyes, and renal concrements from the Lebanese Cenomanian. Swiss Journal of Palaeontology, 140, 15. https://doi.org/10.1186/s13358-021-00229-9

2020

• Pohle, A., Fuchs, D., Korn D. & Klug, C. (2020). Spatial distribution of oncocerid cephalopods on a Middle Devonian bedding plane suggests semelparous life cycle. Scientific Reports, 10, 2847. https://doi.org/10.1038/s41598-020-59507-0
• Frey, L., Pohle, A., Rücklin, M. & Klug, C. (2020). Fossil-Lagerstätten, palaeoecology and preservation of invertebrates and vertebrates from the Devonian in the eastern Anti‐Atlas, Morocco. Lethaia, 53(2), 242-266. https://doi.org/10.1111/let.12354

2019

• Pohle, A., Klug, C., Toom, U. & Kröger, B. (2019). Conch structures, soft-tissue imprints and taphonomy of the Middle Ordovician cephalopod Tragoceras falcatum from Estonia. Fossil Imprint, 75(1), 70–78. https://doi.org/10.2478/if-2019-0006
• Klug, C., Pohle, A. & Korn, D. (2019). Empty cephalopod conchs as substrates for gastropod eggs from the Hangenberg Black Shale (Late Devonian) of the Maïder Basin (Morocco). Fossil Imprint, 75(1), 59–63. https://doi.org/10.2478/if-2019-0004
• Klug, C., Landman, N.H., Fuchs, D., Mapes, R.H., Pohle, A., Guériau, P., Reguer, S. & Hoffmann, R. (2019). Anatomy and evolution of the first Coleoidea in the Carboniferous. Communications Biology, 2, 280. https://doi.org/10.1038/s42003-019-0523-2
• Pohle, A., Klug, C. & Haas, M. (2019). Gigantism, Taphonomy and Palaeoecology of Basiloceras, a new oncocerid genus from the Middle Devonian of the Tafilalt (Morocco). Swiss Journal of Palaeontology, 138(1), 151–162. https://doi.org/10.1007/s13358-019-00182-8

2018

• Klug, C., Pohle, A., Kiel, S. & Kröger, B. (2018). A fossilized marble run: the peculiar taphonomy of Ordovician diploporitan blastozoans from Sweden. Swiss Journal of Palaeontology, 137(2), 405–411. https://doi.org/10.1007/s13358-018-0173-x
• Pohle, A. & Klug, C. (2018). Early and Middle Devonian Cephalopods from Hamar Laghdad (Tafilalt, Morocco) and remarks on epicoles and cameral deposits. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 290(1–3), 203–240. https://doi.org/10.1127/njgpa/2018/0776
• Klug, C., Samankassou, E., Pohle, A., De Baets, K., Franchi, F. & Korn, D. (2018). Oases of biodiversity: Devonian mudmounds at Hamar Laghdad, Morocco. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen, 290(1–3), 9–48. https://doi.org/10.1127/njgpa/2018/0772
• Pohle, A. & Klug, C. (2018). Body size of orthoconic cephalopods from the late Silurian and Devonian of the Anti-Atlas (Morocco). Lethaia, 51(1), 126–148. https://doi.org/10.1111/let.12234

2017

• Klug, C., Frey, L., Pohle, A., De Baets, K. & Korn, D. (2017). Palaeozoic Evolution of Animal Mouthparts. Bulletin of Geosciences, 92(4), 511–524. https://doi.org/10.3140/bull.geosci.l648

Non peer-reviewed articles (including archived preprints)

• Stevens, K., Pohle, A., Hoffmann, R. & Immenhauser, A. (2022). Bayesian inference reveals a complex evolutionary history of belemnites. bioRxiv. https://doi.org/10.1101/2022.08.22.504746
• Barido-Sottani, J., Pohle, A., De Baets, K., Murdock, D. & Warnock, R.C.M. (2022). Putting the F in FBD analyses: tree constraints or morphological data? bioRxiv. https://doi.org/10.1101/2022.07.07.499091
• Klug, C. & Pohle, A. (2018). The eastern Amessoui Syncline - a hotspot for Silurian to Carboniferous cephalopod research. Field Guidebook: 10^th International Symposium of Cephalopods Present and Past, Münstersche Forschungen zur Geologie und Paläontologie, 110, 244–260.
• Klug, C., Bohatý, J., Müller, P. & Pohle, A. (2017). Erstes vollständiges Nautiliden-Gehäuse aus der Lahn-Mulde - paläontologische Auswertung des Schurfs “Löhnberg I”, Lkr. Limburg-Weilburg. hessenARCHÄOLOGIE - Jahrbuch für Archäologie und Paläontologie in Hessen, 2016, 27–29.

Conference contributions

Keynote and invited talks

• Pohle, A. (2023). What is a “nautiloid” cephalopod? A modern perspective. 94. Jahrestagung der Paläontologischen Gesellschaft, Jena, Germany.
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X., & Klug, C. (2022). Phylogeny of early Palaeozoic cephalopods. GSA Connects 2022, Denver (Colorado), USA.
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X., Jell, P. & Klug, C. (2022). Early cephalopod evolution: Insights from new methods and old fossils. 11^th International Symposium on Cephalopods Present and Past, London, United Kingdom.

Regular talks

• Pohle, A., Stevens, K., Hoffmann, R. & Immenhauser, A. (2024). ‘Phylogeochemistry’: can trace elements inform belemnite evolutionary relationships? 68^th Annual Meeting of The Palaeontological Association, Erlangen, Germany.
• Pohle, A., Hoffmann, R., Aubrechtová, M., Seuss, B., Kröger, B., Nützel, A., Stevens, K. & Immenhauser, A. (2024). Illuminating the formation cycle of cephalopod cameral deposits using electron beam microanalysis. 95^th Annual Meeting of the Paläontologische Gesellschaft (PalGes), Warsaw, Poland.
• Pohle, A., Stevens, K., Fuchs, D., Hoffmann, R. & Immenhauser, A. (2023). Fossil coleoid cephalopods: The role of morphological character definitions in phylogenetic inference. 94. Jahrestagung der Paläontologischen Gesellschaft, Jena, Germany.
• Pohle, A., Stevens, K. & Klug, C. (2022). Fossil cephalopod phylogenetics: recent advances, remaining challenges and future perspectives. 20^th Swiss Geoscience Meeting, Lausanne, Switzerland. Awarded as best talk by a young researcher
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Bayesian inference of early Palaeozoic cephalopod evolution. World Congress of Malacology 2022, Munich, Germany.
• Pohle, A., Kröger, B., Warnock, R. C. M., Evans, D. H., Fang, X., Cichowolski, M., Aubrechtová, M., King, A. H. & Klug, C. (2020). Phylogenetic analysis of Cambrian-Ordovician nautiloid cephalopods. 18^th Swiss Geoscience Meeting, Zurich, Switzerland (online).
• Pohle, A., Jell, P.A., Zhen, Y.-Y. & Klug, C. (2019). The forgotten latest Cambrian nautiloid cephalopods from Black Mountain (NW Queensland, Australia). 63^rd Annual Meeting of The Palaeontological Association, Valencia, Spain.
• Pohle, A. & Jell, P.A. (2019). Filling the gap: a forgotten fauna of Cambrian nautiloid cephalopods from Australia and its implications. 17^th Swiss Geoscience Meeting, Fribourg, Switzerland.
• Pohle, A. & Klug, C. (2019). Pairwise clustering of Devonian oncocerids (Cephalopoda) suggests semelparity. 90. Jahrestagung Paläontologische Gesellschaft, Munich, Germany.
• Pohle, A. & Klug, C. (2018). Conch size of Siluro-Devonian orthoconic nautiloids from the Anti-Atlas (Morocco) – food availability as a limiting factor? 10^th International Symposium of Cephalopods Present and Past, Fez, Morocco.
• Pohle, A. & Klug, C. (2017). Body size of orthoconic cephalopods from the late Silurian and Devonian of the Anti-Atlas (Morocco). 15^th Swiss Geoscience Meeting, Davos, Switzerland.

Posters

• Pohle, A. (2023). A persistent misnomer: The curious case of the Paleozoic cephalopod Orthoceras. 94. Jahrestagung der Paläontologischen Gesellschaft, Jena, Germany. Awarded 3^rd place as best poster

Session convener & chair

• Pohle, A., Karapunar, B., Neubauer, T.A. & Skawina, A. (2024). S1: Advances in mollusk paleontology. 95^th Annual Meeting of the Paläontologische Gesellschaft (PalGes), Warsaw, Poland.

Co-authored contributions

• Klug, C., Pohle, A., Fuchs, D. & Hoffmann, R. (2022). Exceptionally preserved cephalopod fossils from conservation deposits. 11^th International Symposium on Cephalopods Present and Past, London, United Kingdom.
• Klug, C., Frey, L., Coates, M., Greif, M., Jobbins, M., Pohle, A., Lagnaoui, A., Bel Haouz, W. & Ginter, M. (2022). Anatomy and phylogeny of a close relative of the chondrichthyan Cladoselache from the Devonian of Morocco. *66^th Annual Meeting of The Palaeontological Association, Cork, Ireland.
• Warnock, R.C.M., Barido-Sottani, J., De Baets, K., Sciré, J., Pohle, A. & Murdock, D. (2022). Evaluating different approaches to handling taxonomic uncertainty using the fossilized birth-death process. Evolution 2022, Cleveland (Ohio), USA.
• Klug, C., Samankassou, E., Pohle, A., Zapalski, M. & Korn, D. (2018). Couscous ai frutti di mare - the spectacular Moroccan mudmound locality Hamar Laghdad and its palaeoecology. 16^th Swiss Geoscience Meeting, Bern, Switzerland.
• Samankassou, E., Klug, C., Pohle, A. & Franchi, F. (2018). Oases of biodiversity: Devonian mudmounds at Hamar Laghdad, Morocco. 20^th International Sedimentological Congress, Québec, Canada.
• Klug, C., Samankassou, E., Pohle, A., Zapalski, M. & Korn, D. (2018). Couscous ai frutti di mare – Early Devonian palaeoecology of the Moroccan mudmound locality Hamar Laghdad. 5^th International Palaeontological Congress, Paris, France.
• Klug, C., Frey, L., Pohle, A., De Baets, K. & Korn, D. (2018). Palaeozoic evolution of cephalopod mouthparts. 10^th International Symposium of Cephalopods Present and Past, Fez, Morocco.
• Klug, C., Frey, L., Pohle, A. & Rücklin, M. (2017). Origin of the Konservatlagerstätten of the southern Maïder (Morocco) and gnathostome preservation. 14^th International Symposium on Early and Lower Vertebrates, Chęciny, Poland.

Data sets

• Barido-Sottani, J., Pohle, A., De Baets, K., Murdock, D. & Warnock, R.C.M. (2023). Supplementary materials for “Putting the F into FBD analysis: tree constraints or morphological data?” Zenodo, https://doi.org/10.5281/zenodo.6884063
• Klug, C., Coates, M., Frey, L., Greif, M., Jobbins, M., Pohle, A., Lagnaoui, A., Haouz, W.B., Ginter, M. CT-stacks of the neurocranium of the Late Devonian chondrichthyan Maghriboselache. MorphoSource, https://www.morphosource.org/projects/000492799
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Additional file 1 of Early cephalopod evolution clarified through Bayesian phylogenetic inference. figshare, https://doi.org/10.6084/m9.figshare.19602439.v1
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Additional file 2 of Early cephalopod evolution clarified through Bayesian phylogenetic inference. figshare, https://doi.org/10.6084/m9.figshare.19602445.v1
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Additional file 3 of Early cephalopod evolution clarified through Bayesian phylogenetic inference. figshare, https://doi.org/10.6084/m9.figshare.19602448.v1
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Additional file 4 of Early cephalopod evolution clarified through Bayesian phylogenetic inference. figshare, https://doi.org/10.6084/m9.figshare.19602451.v1
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Additional file 5 of Early cephalopod evolution clarified through Bayesian phylogenetic inference. figshare, https://doi.org/10.6084/m9.figshare.19602454.v1
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Additional file 6 of Early cephalopod evolution clarified through Bayesian phylogenetic inference. figshare, https://doi.org/10.6084/m9.figshare.19602460.v1
• Pohle, A., Kröger, B., Warnock, R.C.M., King, A.H., Evans, D.H., Aubrechtová, M., Cichowolski, M., Fang, X. & Klug, C. (2022). Additional file 7 of Early cephalopod evolution clarified through Bayesian phylogenetic inference. figshare, https://doi.org/10.6084/m9.figshare.19602445.v1
• Klug, C., Landman, N.H., Fuchs, D., Mapes, R.H., Pohle, A., Gueriau, P., Solenn, R. & Hoffmann, R. (2019). RTI_Gordoniconus_AMNH43264v1.ptm. figshare, https://doi.org/10.6084/m9.figshare.8319983
• Klug, C., Landman, N.H., Fuchs, D., Mapes, R.H., Pohle, A., Gueriau, P., Solenn, R. & Hoffmann, R. (2019). RTI_Gordoniconus_AMNH43264v2.rti. figshare, https://doi.org/10.6084/m9.figshare.8319989

Public talks and seminars

• Pohle, A. (2023). The third dimension: How a forgotten fauna of latest Cambrian cephalopods reveals long-standing misinterpretations. Institut für Geologie und Paläontologie, Universität Münster.
• Pohle, A. (2023). Die Formenvielfalt der frühesten Kopffüsser und ihre Evolution. Mineralien- und Fossilienfreunde Aargau, Vorträge, Restaurant Ochsen, Lupfig, Switzerland (in German).
• Pohle, A. (2022). Die Formenvielfalt der frühesten Kopffüsser und ihre Evolution. Mineralien- und Fossilienfreunde Bern, Vorträge Naturhistorisches Museum Bern, Bern, Switzerland (in German).
• Pohle, A. (2019). Diversity and Palaeobiology of nautiloid cephalopods from the Devonian of the eastern Anti-Atlas (Morocco). Nanjing Institute of Geology and Palaeontology, Nanjing, China.
• Pohle, A. (2019). Mützen, Tüten und Spiralen: Die frühe Evolution der Kopffüsser. Öffentliche Museumsvorträge, Paläontologisches Institut und Museum, Universität Zürich, Switzerland (in German).
• Pohle, A. (2016). Seh‘ ich recht? Merkwürdige Lebensformen im Laufe der Erdgeschichte. Lange Nach der Zürcher Museen, Paläontologisches Institut und Museum, Universität Zürich, Switzerland (in German).
• Pohle, A. (2015). Urtümliche Kopffüsser: Kopfüber durch die Urmeere. Lange Nach der Zürcher Museen, Paläontologisches Institut und Museum, Universität Zürich, Switzerland (in German).

Outreach

• Pohle, A. (2022). Cones, sticks and croissants: Early cephalopod evolution. Nature Communities, Behind the Paper. https://communities.springernature.com/posts/cones-sticks-and-croissants-early-cephalopod-evolution

Media coverage

• April 4, 2024 Several articles on Ticinoteuthis chuchichaeschtli in Swiss media:
  https://www.srf.ch/news/schweiz/entdeckung-der-uni-zuerich-duerfen-wir-vorstellen-die-tintenfischart-chuchichaeschtli (in German)
  https://www.tagesanzeiger.ch/palaeontologie-an-der-uni-zuerich-neu-entdeckte-tessiner-tintenfisch-art-wird-chuchichaeschtli-getauft-375547634858 (in German)
  https://www.swissinfo.ch/eng/science/ticino-octopus-species-christened-chuchich%C3%A4schtli/75111337
• March 19, 2024. Article Spotlight: Plectronoceratids (Cephalopoda) from the latest Cambrian at Black Mountain. Communities @ PeerJ. https://peerj.com/blog/post/115284889009/ (with video interview on Youtube)